Butter and flour two eight inch round metal cake pans. Preheat the oven to 325 F.
Set the buttermilk in a small pot and stir in the poppy seeds. Set the pot over low heat. Once the mixture is warm, remove from heat and set aside.
In the bowl of a stand mixer add the sugar and over it zest the lemons. Rub the zest into the sugar with your fingers to allow the oils to release. Stop once it resembles wet sand.
Add the butter, oil, vanilla and salt to the bowl and affix the paddle attachment. Cream the mixture together until light and airy, about 3-5 minutes at medium speed. Stop the mixer and scrape down the bowl to ensure no bits remain unmixed.
With the mixer on, add the eggs one by one. Once they are all in, turn the speed to medium and beat until you have a creamy mixture.
Sift in the flour and baking powder and with the mixer on, pour in the buttermilk poppy seed mix. Mix to just combine.
Divide the batter evenly between the two pans and smooth the batter so it’s in one layer.
Bake the cakes until a cake tester comes out clean, the tops spring back - about 30-35 minutes. Let cool briefly in the pan then invert onto a cooling rack to completely cool.